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The Bonanza Creek LTER is a member of the U.S. LTER Network which is supported by the National Science Foundation (DEB-1636476) and by the USDA Forest Service, Pacific Northwest Research Station (RJVA-PNW-01-JV-11261952-231). Any opinions, findings, and conclusions or recommendations expressed in this material are those of the author(s) and do not necessarily reflect the views of the National Science Foundation.